The picture Sardine Run was captured by French photographer Greg Lecoeur from Nice in June 2015, following a two-week wait to get the perfect shot.
“Millions of sardines are preyed upon by marine predators such as dolphins, marine birds, sharks, whales, penguins, sailfishes and sea lions,” said Lecoeur.
“The hunt begins with common dolphins that have developed special hunting techniques to create and drive baitballs to the surface. In recent years, probably due to overfishing and climate change, the annual Sardine Run has become more and more unpredictable.”
Lecoeur won a 10-day trip for two to Galapagos with National Geographic Expeditions and two 15-minute image portfolio reviews with National Geographic photo editors.
See more of the winning entries here